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Title: Homepage (E. M. Barrows, research, senior thesis)

Barrows’ Homepage

Senior Theses and Tutorials

Currently, I’m especially interested in having senior-thesis students perform hands-on studies of the biodiversity of Dyke Marsh Wildlife Preserve (DMWP) which is part of the national park called George Washington Memorial Parkway in northern Virginia.  

What skills will you learn?

carefully performing scientific work
analyzing data
curating and identifying a select group of arthropods
improving your scientific-communication ability via writing and oral presenting
improving your understanding of scientific papers.

(My lab gives you much guidance that will improve your work.)

How is this project relevant to your career plans?

This project will teach you excellent skills that can help reach the next step in your career development.   Environmental-biology projects such as this one on DMWP are significant stepping stones to graduate and medical schools and biological and other jobs.  

How is this project relevant to our environment?

This biodiversity project helps us to learn how to monitor and manage DMWP arthropods (insects and their kin) as part of stewardship of the national park, and in turn our Earth.   The National Park Service is charged with maintaining the health of DMWP.   The arthropods are key members of the DMWP ecosystem where they aerate soil, function in the Preserve’s complex food web, pollinate, and recycle nutrients.  

What will you work on?

My lab currently has a large, rare sample of DMWP arthropods awaiting senior-thesis students who can process and analyze them.   There are about 20 scientist-years of work left to do on the analysis.   The samples are preserved and ready for work.   Students and I determine which arthropods live in DMWP, which habitats they use, and their abundances, sex ratios, and seasonal cycles in view of conserving these valuable animals.   So far students and I have published three scientific papers on DMWP arthropods: soil arthropods in the low forest, alderflies (Sialidae) and fireflies (Lampyridae) in the forest, open marsh, and forest-marsh ecotone. We are working on manuscripts concerning apid bees (Apidae), damselflies and dragonflies (Odonata), and deer and horse flies (Tabanidae).   There are many more papers to produce regarding our DMWP project.  

How do you learn more about DMWP?

You can start by clicking here to learn more about DMWP on this Website.

There is room for several seniors and undergraduate tutorial-research students in my lab.   If you are interested in working my lab on a senior-thesis project, please contact me at barrowse@georgetown.edu.   If you are interested in GUROP support in my Lab, please check out the GUROP information on GU’s Website and contact me.


I’ve been the research advisor for many senior-theses and other undergraduates in my Lab (list below).   Seven of these students coauthored peer-reviewed papers, or book chapters, with me so far (each indicated with an asterisk).   Several students gave reports at scientific meetings outside Georgetown Campus (indicated with **), and two won awards for excellent theses (indicated with ***).   Ms. Allison Beck co-published her thesis with Dr. Conrad Labandeira (Smithsonian Institution).   An undergraduate Mr. Florian Schiestl (University of Vienna, Austria), who was not a senior-thesis student, came to work in my Laboratory in fall semester 1994 on bees.   We published a paper together.

Senior Theses

1. Michael Francis Kane. 1976. Biological observations on the Japanese Beetle, Popillia japonica (Newman) (Coleoptera: Scarabaeidae). Laboratory thesis.

2. Mary-Louise A. Tracy. 1976. Periodicity in mating behavior of females of Aphelinus varipes (Forester) Hymenoptera: Chalcidoidea: Aphelinidae). Laboratory thesis.

3. Jean Basior. 1976. Chalcidoid wasp parasitizing behavior. Library thesis.

4. Anthony P. Acquavella*. 1977. Feeding and other behavior in captive, juvenile Mockingbirds, Mimus polyglottos (Passeriformes: Mimidae). Laboratory thesis.

5. Patrick J. Lanigan*. 1977. Sexual behavior of Murgantia histrionica (Hahn) (Hemiptera: Pentatomidae) and some other aspects of its biology. Laboratory thesis.

6. Laurie A. Heffernan. 1978. Insect pollinators and flower perforators of Ocimum basilicum (Methaceae). Laboratory thesis.

7. Roland M. Marshal, III. 1978. Defense mechanisms of arthropods. Library thesis.

8. Ronald de Andrade. 1979. The distribution and biology of Washington, D.C., harvestmen. Library thesis.

9. Thomas M. Hughes. 1979. The biology of agelenid spiders. Library thesis.

10. John J. Lesica. 1979. Konrad Lorenz: Ethological contributions to animal behavior. Library thesis.

11. David P. Norton. 1979. A study of bumble bees (genus Bombus) including experimental observation of pollination of Actinomeris by male Bombus impatiens. Laboratory thesis.

12. Scott R. Schoem. 1979. Cannibalism and feeding behavior in young Stagmomantis carolina (Orthoptera: Mantidae). Laboratory thesis.

13. Andrea Canary. 1980. Crops and pollination in the Glover Park Community Vegetable Garden. Laboratory thesis.

14. James S. DeFilippo*. 1980. Urban gardener knowledge of arthropods. Laboratory thesis.

15. Walter J. Giblin. 1980. Soil arthropods in an urban community vegetable garden. Laboratory thesis.

16. Timothy O'Malley. 1980. Body language with respect to age and sex in a dental clinic waiting room. Laboratory thesis.

17. Morad Tavallale*. 1980. Urban gardener knowledge of arthropods. Laboratory thesis.

18. Diane B. Hobbs. 1981. Offspring number and birth behavior in the Black Bean Aphid, Aphis fabae (Homoptera: Aphididae). Laboratory thesis.

19. Antonio E. Rabassa. 1981. Prey size and feeding behavior of the Great Barracuda (Sphyraena barracuda) and its consequence to the bathers. Library thesis.

20. John Riggs. 1981. Karl von Frisch's visual sense studies with the Honey Bee, Apis melifera. Library thesis.

21. Mark J. Uggeri. 1981. Aspects of the biology of aphidophagous Syrphidae (Diptera). Library thesis.

22. German Newall. 1982. Carpenter bee anatomy.

23. Louis S. Angioletti. 1982. Pheromones are a potential alternative to conventional pesticides for controlling lepidopteran pests.

24. James Chun. 1982. Proposed mechanisms of homing behavior in pigeons. Library thesis.

25. John V. Grieco. 1982. Insecticides as a method of integrated control of the Tsetse Fly. Library thesis.

26. Mary Macnamara. 1982. A survey of public knowledge of Dog behavior and care. Laboratory thesis.

27. Salvatore M. Moffa. 1982. Diseases of Apis mellifera. Library thesis.

28. Joseph Randell. 1982. Mate choice in scorpionflies. Library thesis.

29. Mohammad R. Momtaz. 1983. The role of aggression in the reproductive behavior of confined troops of Japanese Macaques. Laboratory thesis.

30. Brian V. Reamy. 1983. A survey of attitudes and knowledge concerning insects and pesticides in students and the general public. Laboratory thesis.

31. Michael A. Sergi. 1983. A survey of knowledge and attitudes about insects in selected groups of people. Laboratory thesis.

32. David C. Weyn. 1983. Seasonal size difference on Chinese Mantids, Tenodera ardifolia sinensis (Orthoptera: Mantidae). Laboratory thesis.

33. Mary E. Pieschel. 1984. Possible factors affecting female Mexican Bean Beetle, Epilachna varivestis, fecundity. Laboratory thesis.

34. Joanne M. Fugii. 1985. The hawk-dove game: A computer demonstration. Laboratory thesis.

35. Suzanne M. Girard. 1985. Monandry, serial polyandry, size, season, fertility, and fecundity in female Mexican Bean Beetles, Epilachna varivestis (Coleoptera: Coccinellidae). Laboratory thesis.

36. Mary K. Landers. 1985. Possible factors affecting female fecundity and fertility in the Mexican Bean Beetle, Epilachna varivestis. Laboratory thesis.

37. Martin S. O'Brien. 1985. The correlation of six independent factors in the fecundity and fertility of female Mexican Bean Beetle, Epilachna varivestis. Laboratory thesis.

38. Jeanne M. Reamy. 1985. The effect of six independent variables on the fecundity and fertility of female Mexican Bean Beetles, Epilachna varivestis. Laboratory thesis.

39. Daniel P. O'Day. 1986. Mating regime, size, mate size, season, fertility, and fecundity in female Mexican Bean Beetles, Epilachna varivestis (Coleoptera: Coccinellidae). Laboratory thesis.

40. Gregory M. Pitaro. 1986. Mating regime, dry weight, femur length, season, fertility, and fecundity in female Mexican Bean Beetles, Epilachna varivestis. (Coleoptera: Coccinellidae). Laboratory thesis.

41. Fitzpatric C. Wilson. 1987. Definitions and insight into altruism and related terms. Library thesis.

42. Ronald S. Guibord. 1988. Courtship and copulatory behavior in experienced and virgin male Mexican Bean Beetles, Epilachna varivestis (Coleoptera: Coccinellidae). Laboratory thesis.

43. Marie Mancini. 1988. Courtship and copulatory behavior of once-copulation-experienced males and virgin males of Mexican Bean Beetles, Epilachna varivestis . (Coleoptera: Coccinellidae). Laboratory thesis.

44. Matthew P. Smith* 1989. Effects of larval case size, host plant species, and ambient temperature on case internal temperatures in the bagworm, Thyridopteryx ephemeraeformis (Lepidoptera: Psychidae).

45. Edward L. Mills. 1990. Effects of height zone and host condition of Arbor Vitae (Thuja occidentalis) on the Bagworm Thyridopteryx ephemeraeformis (Lepidoptera: Psychidae)

46. Renee S. Webbink. 1990. Mating behavior of experienced and inexperienced male Mexican Bean Beetles (Epilachna varivestis). Laboratory thesis.

47. George Aguiar. 1991. Ecological competition: Ant interactions at bonanza food sources (Hymenoptera: Formicidae). Laboratory thesis.

48. Ann H. Partridge. 1991. Ecological competition: Ant interactions. Laboratory thesis.

49. Joel A. Wilsnack. 1991. Ecological competition: Ant interactions (Hymenoptera: Formicidae). Laboratory thesis.

50. Lynne Barre. 1992. The late-season mimic-scarcity hypothesis: A test with yellowjackets and flower fly mimics in a northern broadleaf forest. Laboratory thesis.

51. Wanda G. Noguera-Irizzary. 1992. The late-season mimic-scarcity hypothesis: A test with black-and-white yellowjackets and their flower fly mimics in a northern broadleaf forest. Laboratory thesis.

52. Samantha S. Wolf* 1992. Yellowjacket abundance, distribution, and flight periods in a Northern broadleaf forest (Hymenoptera: Vespidae). Laboratory thesis.

53. Darren M. Lynch*. 1993. Diflubenzuron effects on non-target vespid wasps in a northern broadleaf forest (Hymenoptera: Vespidae). Laboratory thesis.

54. Holi D. Ieng. 1994. Trichodezia albovittata in a central Appalachian broadleaf forest (Lepidoptera: Geometridae): Flight periods, flight areas, and diflubenzuron. Laboratory thesis.

55. Cheryl R. Ladota. 1994. To survive and multiply. The role of zoos in saving endangered and threatened species through captive breeding. Library thesis.

56. Angela L. Reed. 1994. The effect of diflubenzuron on beneficial insects: A study of vespid wasps in a central Applachian broadleaf forest (Hymenoptera:Vespidae)

57. Ali Al-Atar. 1995. Asilids in a central Appalachian broadleaf forest, West Virginia: A study of genera, flight periods, abundances, diflubenzuron, and the late-season-mimic-scarcity hypothesis. Laboratory thesis.

58. Melissa R. Reilly. 1995. Asilids in the Fernow Experimental Forest of West Virginia: A study of genera, flight periods, abundances, diflubenzuron and the late-season- mimic-scarcity hypothesis. Laboratory thesis.

59. Philip J. Wellington 1995. Asilids in a central Appalachian broadleaf forest: Genera, flight periods, diflubenzuron and the late-season-mimic-scarcity hypothesis. Laboratory thesis.

60. James V. Tedesco. 1995. Ceropales maculata fraterna in a central Appalachian broadleaf forest (Hymenoptera: Pompilidae): Flight areas, flight periods, pesticide effects, and mimicry.

61. Mark V. Siveri. 1995. Selected vespid and sphecid wasps in a central Appalachian broadleaf forest (Hymenoptera: Vespidae and Sphecidae): Abundances, flight periods and areas, diflubenzuron, and mimicry. Laboratory thesis.

62. Gabriela E. Zöllner***. 1995. Worker size of the Eastern Yellowjacket (Vespula maculifrons) in different years and time of the flight season in a central Appalachian broadleaf forest. Laboratory thesis.

63. Gurpal S. Ahluwalia. 1996. Selected Asilidae of a central Appalachian broadleaf forest: Species composition, sex ratios, flight periods, and possible trap-site effect and diflubenzuron. Laboratory thesis.

64. Brice C. Bartley. 1996. Lightningbugs (Coleoptera: Lampyridae) of a central Appalachian broadleaf forest: Genus composition, flight periods, sex ratios, trap-site effect, and diflubenzuron. Laboratory thesis.

65. Allison L. Beck**. 1996. Plant-insect interactions: Survey of the Early Permian Period with implications for the environmental saturation hypotheses and expanding resources hypothesis. (Co-advised with Conrad Labandeira, Smithsonian Institution) Laboratory thesis. (Allison published her thesis as: Beck, A. L. and C. C. Labandeira. 1998. Early Permian insect folivory on a gigantopterid-dominated riparian flora from north-central Texa. Palaeogeography, Palaeoclimatology, Palaeoecology 142 139173.)

66. Anthony A. Marchese. 1996. Lightningbugs (Coleoptera: Lampyridae) in a central Appalachian broadleaf forest: Genera present, sex ratio, flight periods, and possible diflubenzuron and trap-site effects. Laboratory thesis.

67. Matthew D. Chua. 1997. Tidal Freshwater Marshes and the Need for Their Management. Library thesis with some original data.

68. Jason Co. 1997. Evidence supporting dinosaur-extinction hypotheses points toward multiple causality for the extinction at the Cretaceous-Tertiary (K/T) boundary, and lends insight into hypotheses regarding periodic mass extinction. Library thesis.

69. Dollie D. Silpasuvan. 1997. The evolution of menstruation and endometrial cycles. Library thesis.

70. Brian N. Smith. 1999. Flower Flies (Diptera: Syrphidae), Biodiversity, and Edge Efect, in Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis. 39 pp.

71. Christopher Watson. 1999. Bee (Hymenoptera: Apiformes) Diversity, and a Search for an Edge Effect, in the Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis.

72. Ho V. Dzung**. 2000. Bee Diversity and Habitat Effects in Dyke Marsh Wildlife Preserve, Virginia.

73. Andrew W. Hung**. 2000. Diversity, Flight Periods, and Habitat Effects of Noctuid Moths (Lepidoptera: Noctuidae) in Dyke Marsh Wildlife Preserve (DMWP), Virginia. Laboratory thesis.

74. John W. Kraemer**. 2000. Habitat and Flower Fly (Diptera: Syrphidae) Biodiversity and Flight Periods in Dyke Marsh Wildife Preserve (DMWP), Virginia. Laboratory thesis.

75. Christopher P. McManus**. 2000. Putative Batesian Mimicry in the Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis. Laboratory thesis.

76. Michael Gwinn. 2001. Fly Harrassment of Wild Ponies on Assateague Island, Maryland. (with a researcher at the National Zoological. Laboratory thesis.

77. Keith Post. 2002. Flower Fly (Diptera: Syrphidae) Biodiversity, Habitat Uses and Flight Periods in Dyke Marsh Wildlife Preserve (DMWP), Virginia. Laboratory thesis.

78. Anne M. McIntyre*. 2003. Alderfly (Neuroptera: Sialidae) Flight Periods and Habitat Uses in Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis.

79. Alexander T. Taft, III. 2003. Odonata Abundance, Habitat, and Flight Periods in the Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis.

80. Shawna B. Arsenault*. 2004. Firefly (Coleoptera: Lampyridae) Biodiversity, Habitat Use, and Flight Periods in Dyke Marsh Wildlife Preserve (DMWP), Virginia. Laboratory thesis.

81. Daniel G. Balogh. 2005. Ichneumonid (Hymenoptera: Ichneumonidae) Flight Periods, Abundances, and Habitat Distribution in Dyke Marsh Preserve, Virginia. Laboratory thesis.

82. Christianne R. Bird. 2005. Ichneumon Wasps (Hymenoptera: Ichneumonidae) Flight Periods, Abundance, and Habitat Preferences in a National Park. Laboratory thesis.

83. Marie LaRiviere. 2005. Feral Cats and Marsupials in Australia. Library thesis.

84. Ann Bobel. 2006. Ichneumon Wasps (Hymenoptera: Ichneumonidae) Flight Periods, Abundance, and Habitat Preferences in a National Park. Laboratory thesis.

85. Bao Q. Chung. 2006. Soil Mites in a National Park. Laboratory thesis.

86. Claire E. McNulty. 2007. Odonate (Odonata: Anisoptera, Zygoptera) Habitat Use, Flight Periods, and Sex Ratios in Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis.

87. Daniel F. Vance. 2008. Tabanid (Diptera: Tabanidae) Abundance, Flight, Seasonality, Habitat Distribution, and Sex Ratios in the Dyke Marsh Wildlife Preserve, Virginia. Laboratory thesis.

88. Michael E. Kolleng. 2009. Apid Bee (Hymenoptera: Apidae) Flight Periods, Sex-Ratios, Abundance, and Habitat Distribution in a Mid-Atlantic National Park. Laboratory thesis.

89. George A. Roche. 2010. Augochlorine Bee Populations, Distribution, and Biodiversity within a Mid-Atlantic National Park. Laboratory thesis.

90. Maria Baquerizo**. 2011. Lasioglossum Bee Populations, Distribution, and Biodiversity within a Mid-Atlantic National Park. RISE Project. Poster presented at the 2011 annual meeting of the Ecological Society of America, Austin, TX.

91. Harrison A. Bookstein**. 2012. Pollinium Acquisition in Three Pollinator Taxa of Asclepias syriaca, Common Milkweed (Apocynaceae). RISE Project.

(updated April 2012)



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